Marengo
Transfers of genera to other families - APGIII
Recent work on the phylogeny of plant families has led to some significant revisions of family boundaries, with a number of genera being moved
from one family to another, and some families being assimilated within other families. Far more detail is available at the website of the Angiosperm Phylogeny Group
While we try to keep our plant list taxonomy as up-to-date as
possible at generic, specific and subspecific level, we have not leapt fully into the new familial parameters.
In order to provide plant list users with as much information as possible, the following are changes that would apply to families and genera currently included
in our Cypriot, Greek, Macaronesian, Dolomites, Spanish and Pyrenean plant lists. The list below is of families with altered parameters, and genera which have been assigned to them.
s.l. (sensu lato) refers to families that still exist after the changes, albeit in a more restricted sense.
It is important to note that individual changes have varying degrees of support and it is likely that some will eventually be adopted, while others will
not. Those in double closed square brackets [[]], have been recently considered as separate families, but the current concensus is towards (re-)inclusion.
October 2009.
Just when we thought it was safe to go back into the taxonomic waters, the changes of APG111 appeared in the Botanical Journal of the Linnaean Society.
The broad changes are in red below.
ADOXACEAE (also includes Adoxa)
Sambucus (ex-Caprifoliaceae s.l.)
Viburnum (ex-Caprifoliaceae s.l.)
AGAPANTHACEAE
October 2009. Agapanthaceae is transferred into an expanded Amaryllidaceae s.l.
AGAVACEAE
October 2009. Agavaceae is transferred into an expanded Asparagaceae s.l.
ALLIACEAE
Alliaceae is transferred into an expanded Amaryllidaceae
ALTINGIACEAE
Liquidamber (ex-Hamamelidaceae s.l.)
ALSTROEMERIACEAE
Alstroemeria (ex-Liliaceae)
AMARANTHACEAE
Arthrocnemum (ex-Chenopodiaceae)
Atriplex (ex-Chenopodiaceae)
Bassia (ex-Chenopodiaceae)
Beta (ex-Chenopodiaceae)
Camphorosma (ex-Chenopodiaceae)
Chenoleoides (ex-Chenopodiaceae)
Chenopodium (ex-Chenopodiaceae)
Corispermum (ex-Chenopodiaceae)
Einadia (ex-Chenopodiaceae)
Hablitzia (ex-Chenopodiaceae)
Halocnemum (ex-Chenopodiaceae)
Kochia (ex-Chenopodiaceae)
Noaea (ex-Chenopodiaceae)
Petrosimonia (ex-Chenopodiaceae)
Polycnemum (ex-Chenopodiaceae)
Salsola (ex-Chenopodiaceae)
Sarcocornia (ex-Chenopodiaceae)
Spinacia (ex-Chenopodiaceae)
Suaeda (ex-Chenopodiaceae)
Traganum (ex-Chenopodiaceae)
AMARYLLIDACEAE (also includes Ipheion)
October 2009. Amaryllidaceae is expanded to include Alliaceae and Agapanthaceae
Acis (ex-Amaryllidaceae s.str.)
Agapanthus (ex-Liliaceae s.l., ex-Agapanthaceae)
Allium (ex-Liliaceae s.l., ex-Alliaceae)
Amaryllis (ex-Amaryllidaceae s.str.)
Crinum (ex-Amaryllidaceae s.str.)
Galanthus (ex-Amaryllidaceae s.str.)
Leucojum (ex-Amaryllidaceae s.str.)
Narcissus (ex-Amaryllidaceae s.str.)
Nerine (ex-Amaryllidaceae s.str.)
Nothoscordum (ex-Liliaceae s.l., ex-Alliaceae)
Pancratium (ex-Amaryllidaceae s.str.)
Sternbergia (ex-Amaryllidaceae s.str.)
APHYLLANTHACEAE
October 2009. Aphyllanthaceae is transferred into an expanded Asparagaceae s.l.
APOCYNACEAE
Araujia (ex-Asclepiadaceae)
Asclepias (ex-Asclepiadaceae)
Caralluma (ex-Asclepiadaceae)
Ceropegia (ex-Asclepiadaceae)
Cionura (ex-Asclepiadaceae)
Cynanchum (ex-Asclepiadaceae)
Periploca (ex-Asclepiadaceae)
Vincetoxicum (ex-Asclepiadaceae)
ARACEAE
Lemna (ex-Lemnaceae)
ARALIACEAE
Hydrocotyle (ex-Umbelliferae s.l.)
ASPARAGACEAE
October 2009. Asparagaceae is expanded to include Agavaceae, Aphyllanthaceae, Hyacinthaceae and Ruscaceae
Agave (ex-Agavaceae s.l.)
Anthericum (ex-Liliaceae s.l., ex-Agavaceae s.l.)
Aphyllanthes (ex-Liliaceae s.l., ex-Aphyllanthaceae)
Asparagus (ex-Liliaceae s.l.)
Bellevalia (ex-Liliaceae s.l., ex-Hyacinthaceae)
Brimeura (ex-Liliaceae s.l., ex-Hyacinthaceae)
Charybdis (ex-Liliaceae s.l., ex-Hyacinthaceae)
Chlorophytum (ex-Liliaceae s.l., ex-Agavaceae s.l.)
Convallaria (ex-Liliaceae s.l., ex-Ruscaceae)
Dipcadi (ex-Liliaceae s.l., ex-Hyacinthaceae)
Dracaena (ex-Agavaceae s.l., ex-Ruscaceae)
Drimia (ex-Liliaceae s.l., ex-Hyacinthaceae)
Furcraea (ex-Agavaceae s.l., ex-Agavaceae s.l.)
Hyacinthella (ex-Liliaceae s.l., ex-Hyacinthaceae)
Hyacinthus (ex-Liliaceae s.l., ex-Hyacinthaceae)
Maianthemum (ex-Liliaceae s.l., ex-Ruscaceae)
Muscari (ex-Liliaceae s.l., ex-Hyacinthaceae)
Ornithogalum (ex-Liliaceae s.l., ex-Hyacinthaceae)
Othocallis (ex-Liliaceae s.l., ex-Hyacinthaceae)
Paradisea (ex-Liliaceae s.l., ex-Agavaceae s.l.)
Polygonatum (ex-Liliaceae s.l., ex-Ruscaceae)
Prospero (ex-Liliaceae s.l., ex-Hyacinthaceae)
Ruscus (ex-Liliaceae s.l., ex-Ruscaceae)
Sanseviera (ex-Liliaceae s.l., ex-Ruscaceae)
Schnarfia (ex-Liliaceae s.l., ex-Hyacinthaceae)
Scilla (ex-Liliaceae s.l., ex-Hyacinthaceae)
Semele (ex-Liliaceae s.l., ex-Hemerocallidaceae)
ASPHODELACEAE
October 2009. Asphodelaceae is transferred into Xanthorrhoeaceae
BORAGINACEAE Either Cordiaceae, Heliotropaceae, Hydrophyllaceae, etc are all separated from Boraginaceae, or are all included.
Cordia (ex-Cordiaceae)
Phacelia (ex-Hydrophyllaceae)
Wigandia (ex-Hydrophyllaceae)
CALCEOLARIACEAE
Calceolaria (ex-Scrophulariaceae s.l.)
CANNABACEAE
Celtis (ex-Ulmaceae s.l.)
CAPRIFOLIACEAE
October 2009. Caprifoliaceae is expanded to include Dipsacaceae, Linnaeaceae, Morinaceae, Valerianaceae
The alternative is the existence of a number of smaller families in order to maintain Dipsacaceae and Valerianaceae relatively intact.
CELASTRACEAE
Parnassia (ex-Parnassiaceae)
CLEOMACEAE
Cleome (ex-Capparaceae s.l.)
COLCHICACEAE (also includes Gloriosa)
Androcymbium (ex-Liliaceae s.l.)
Colchicum (ex-Liliaceae s.l.)
Merendera (ex-Liliaceae s.l.)
[[ CORDIACEAE
Either Cordiaceae, Heliotropaceae, Hydrophyllaceae, etc are all separated from Boraginaceae, or are all included.]]
CYMODOCEACEAE
Cymodocea (ex-Zannichelliaceae)
CYTINACEAE
Cytinus (ex-Rafflesiaceae s.l.)
DROSOPHYLLACEAE
Drosophyllum (ex-Droseraceae)
ERICACEAE
Empetrum (ex-Empetraceae)
Hypopitys (ex-Monotropaceae)
Moneses (ex-Pyrolaceae)
Monotropa (ex-Monotropaceae)
Orthilia (ex-Pyrolaceae)
Pyrola (ex-Pyrolaceae)
GROSSULARIACEAE
Ribes (ex-Saxifragaceae s.l.)
[[ HELIOTROPIACEAE
Either Cordiaceae, Heliotropaceae, Hydrophyllaceae, etc are all separated from Boraginaceae, or are all included.
Heliotropium (ex-Boraginaceae) - includes Ceballosia ]]
HEMEROCALLIDACEAE
October 2009. Hemerocallidaceae is transferred into Xanthorrhoeaceae s.l.
HYACINTHACEAE
October 2009. Hyacinthaceae is transferred into an expanded Asparagaceae s.l.
HYDROCHARITACEAE
Najas (ex-Najadaceae)
[[ HYDROPHYLLACEAE
Either Cordiaceae, Heliotropaceae, Hydrophyllaceae, etc are all separated from Boraginaceae, or are all included.]]
HYPERICACEAE
Hypericum (ex-Guttiferae)
LABIATAE (LAMIACEAE)
Vitex (ex-Verbenaceae)
LILIACEAE sensu stricto
Erythronium (ex-Liliaceae s.l.)
Fritillaria (ex-Liliaceae s.l.)
Gagea (ex-Liliaceae s.l.)
Lilium (ex-Liliaceae s.l.)
Lloydia (ex-Liliaceae s.l.)
Streptopus (ex-Liliaceae s.l.)
Tulipa (ex-Liliaceae s.l.)
LINDERNIACEAE
Lindernia (ex-Scrophulariaceae s.l.)
LINNAEACEAE (also includes Kolkwitzia)
Abelia (ex-Caprifoliaceae s.l.)
Linnaea (ex-Caprifoliaceae s.l.)
MALVACEAE (also includes Adansonia, Dombeya, Fremontodendron, Sterculia, Theobroma)
Tilia (ex-Tiliaceae)
MELANTHIACEAE
Paris (ex-Liliaceae s.l.)
Veratrum (ex-Liliaceae s.l.)
MONTIACEAE
Montia (ex-Portulacaceae)
MYRSINACEAE
October 2009. Included in Primulaceae.
NARTHECIACEAE
Narthecium (ex-Liliaceae s.l.)
NITRARIACEAE
Peganum (ex-Zygophyllaceae)
OROBANCHACEAE
Bartsia (ex-Scrophulariaceae s.l.)
Euphrasia (ex-Scrophulariaceae s.l.)
Lathraea (ex-Scrophulariaceae s.l.)
Lesquereuxia (ex-Scrophulariaceae s.l.)
Melampyrum (ex-Scrophulariaceae s.l.)
Odontites (ex-Scrophulariaceae s.l.)
Orobanche
Parentucellia (ex-Scrophulariaceae s.l.)
Pedicularis (ex-Scrophulariaceae s.l.)
Phelypaea (ex-Scrophulariaceae s.l.)
Rhinanthus (ex-Scrophulariaceae s.l.)
Tozzia (ex-Scrophulariaceae s.l.)
PAPAVERACEAE
Ceratocapnos (ex-Fumariaceae)
Corydalis (ex-Fumariaceae)
Fumaria (ex-Fumariaceae)
Hypecoum (ex-Fumariaceae)
Platycapnos (ex-Fumariaceae)
Rupicapnos (ex-Fumariaceae)
Sarcocapnos (ex-Fumariaceae)
PASSIFLORACEAE
Turnera (ex-Turneraceae)
PENTAPHYLACACEAE
Visnea (ex-Theaceae)
PHRYMACEAE
Mimulus (ex-Scrophulariaceae s.l.)
PLANTAGINACEAE
Antirrhinum (ex-Scrophulariaceae s.l.)
Callitriche (ex-Callitrichaceae s.l.)
Campylanthus (ex-Scrophulariaceae s.l.)
Chaenorhinum (ex-Scrophulariaceae s.l.)
Cymbalaria (ex-Scrophulariaceae s.l.)
Digitalis (ex-Scrophulariaceae s.l.)
Erinus (ex-Scrophulariaceae s.l.)
Globularia (ex-Globulariaceae s.l.)
Gratiola (ex-Scrophulariaceae s.l.)
Hebe (ex-Scrophulariaceae s.l.)
Hippuris (ex-Hippuridaceae s.l.)
Isoplexis (ex-Scrophulariaceae s.l.)
Kickxia (ex-Scrophulariaceae s.l.)
Linaria (ex-Scrophulariaceae s.l.)
Littorella (ex-Scrophulariaceae s.l.)
Maurandya (ex-Scrophulariaceae s.l.)
Misopates (ex-Scrophulariaceae s.l.)
Nemesia (ex-Scrophulariaceae s.l.)
Parahebe (ex-Scrophulariaceae s.l.)
Penstemon (ex-Scrophulariaceae s.l.)
Plantago
Scoparia (ex-Scrophulariaceae s.l.)
Sibthorpia (ex-Scrophulariaceae s.l.)
Veronica (ex-Scrophulariaceae s.l.)
POSIDONIACEAE
Posidonia (ex-Potamogetonaceae s.l.)
POTAMOGETONACEAE
Zannichellia (ex-Zannichelliaceae)
PRIMULACEAE
Heberdenia (ex-Myrsinaceae)
Pleiomeris (ex-Myrsinaceae)
RUBIACEAE
Theligonum (ex-Theligonaceae)
RUPPIACEAE
Ruppia (ex-Potamogetonaceae s.l.)
RUSCACEAE
October 2009. Ruscaceae is transferred into an expanded Asparagaceae s.l.
RUTACEAE
Cneorum (ex-Cneoraceae)
Neochamaelea (ex-Cneoraceae)
SANTALACEAE
Arceuthobium (ex-Loranthaceae s.l.)
Viscum (ex-Loranthaceae s.l., ex-Viscaceae)
However, Nickrent (2010) [pdf] suggests that Viscaceae (Arceuthobium, Viscum) should be separated from Santalaceae, as should Thesiaceae (Kunkelliaceae, Thesium).
SAPINDACEAE
Acer (ex-Aceraceae)
Aesculus (ex-Hippocastanaceae)
SCROPHULARIACEAE
Buddleja (ex-Loganiaceae, ex-Buddlejaceae)
Diascia (ex-Scrophulariaceae s.l.)
Limosella (ex-Scrophulariaceae s.l.)
Myoporum (ex-Myoporaceae)
Scrophularia (ex-Scrophulariaceae s.l.)
Sutera (ex-Scrophulariaceae s.l.)
Verbascum (ex-Scrophulariaceae s.l.)
SMILACACEAE
Smilax (ex-Liliaceae s.l.)
TALINACEAE
Talinum (ex-Portulacaceae)
THEOPHRASTACEAE
October 2009. The genus Samolus is returned to Primulaceae.
TOFIELDIACEAE
Tofieldia (ex-Liliaceae s.l.)
TYPHACEAE
Sparganium (ex-Sparganiaceae s.l.)
XANTHORRHOEACEAE
October 2009. Xanthorrhoeaceae is expanded to include Asphodelaceae and Hemerocallidaceae.
Aloe (ex-Liliaceae s.l.)
Asphodeline (ex-Liliaceae s.l., ex-Asphodelaceae)
Asphodelus (ex-Liliaceae s.l., ex-Asphodelaceae)
Eremurus (ex-Liliaceae s.l., ex-Asphodelaceae)
Hemerocallis (ex-Liliaceae s.l., ex-Hemerocallidaceae)
Phormium (ex-Liliaceae s.l., ex-Hemerocallidaceae)
Simethis(ex-Liliaceae s.l., ex-Hemerocallidaceae)
ZOSTERACEAE
Zostera (ex-Potamogetonaceae s.l.)
Lance Chilton and Marengo 2012
Marengo, 17 Bernard Crescent, Hunstanton PE36 6ER, England